In a city park, Surly the Squirrel has finally gone too far with his latest caper leading to the animal community's winter food cache being destroyed. Now exiled, Surly and his rat buddy Buddy's collective nightmare on the streets ends when they discover a nut store to raid. Meanwhile, the squirrels, the heroic Andie and the ditsy Grayson, are charged by Raccoon to find a new food source and Andie runs into Surly. With no other options, she arranges a deal to help in Surly's heist for the colony, even while Surly fully intends to betray it. However, there is more going on with the nut store being a front for bank robbers while Raccoon has his own agenda to ensure his own power. In the mayhem to come, Surly finds himself challenged in ways he never expected and discovering the real prize to treasure in this adventure.